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    • The Margaret Formation is a geologic formation of the Eureka Sound Group in the Sverdrup Basin in Northwest Territories and Nunavut, Canada. The unit belonging to the Eureka Sound Group which crops out at Ellesmere Island preserves fossils dating back to the Early Eocene period, or Wasatchian in the NALMA classification.
      The Margaret Formation comprises sandstones, sandy siltstones, clay-rich sandstones, coal seams and clay-rich coal seams and volcanic ash beds. The thickness of the formation, which overlies the Mount Moore Formation, reaches about 140 metres (460 ft). Radiometric dating of the formation provided ages of 52.6 ± 1.9 Ma (2010) and 53.7 ± 0.6 Ma (2017).
      The area where the formation was deposited in the Early Eocene experienced a much warmer climate than the High Arctic today, with mean annual temperatures ranging from 7.6 to 12.9 °C (45.7 to 55.2 °F) and warmest month mean temperatures from 18.2 to 22.2 °C (64.8 to 72.0 °F). The deltaic to swamp environment of the Margaret Formation has provided a diverse fauna of various groups of mammals, birds (Presbyornis and Gastornis), reptiles (turtles, snakes, lizards and crocodiles) and fish.


      Description


      The Margaret Formation comprises sandstones, sandy siltstones, clay-rich sandstones, coal seams and clay-rich coal seams and volcanic ash beds. The thickness of the formation, which overlies the Mount Moore Formation, reaches about 140 metres (460 ft).
      The formation was probably deposited in a lush proximal delta front to delta plain environment, with abundant channels and coal swamps.


      = Dating

      =
      A volcanic ash layer containing crandallite in the middle of the formation was dated using U-Pb radiometric dating in 2017 to 53.7 ± 0.6 Ma. In 2010, ashes of the formation were dated to 52.6 ± 1.9 Ma.


      = Arctic climate of the Early Eocene

      =

      During the Early Eocene, the climate of much of northern North America was warm and wet, with mean annual temperatures (MAT) as high as 20 °C (68 °F), mean annual precipitation (MAP) of 100 to 150 centimetres (39 to 59 in), mild frost-free winters (coldest month mean temperature >5 °C (41 °F)), and climatic conditions that supported extensive temperate forest ecosystems.
      Ensemble estimates of mean annual temperatures for the high-latitude fossil localities in Arctic Canada ranged from 7.6 to 12.9 °C (45.7 to 55.2 °F), with the range of coldest month mean temperature from 1.3 to 4.2 °C (34.3 to 39.6 °F) and warmest month mean temperatures from 18.2 to 22.2 °C (64.8 to 72.0 °F). Mean annual precipitation estimates for the Margaret Formation ranged between 131 and 180 centimetres (52 and 71 in). The mean summer precipitation has been estimated at 1,134 millimetres (44.6 in) and mean winter precipitation at 366 millimetres (14.4 in).
      The fossils and sedimentology indicate a lush, rain forest community on a coastal delta plain. Multiple palaeoclimate proxies, ranging from oxygen isotope analysis of vertebrate
      bones and teeth to palaeofloral analyses, estimate a mild temperate climate for the Eocene High Arctic, where winters remained at or just above freezing and summer temperatures extended to 20 °C (68 °F) or higher. These temperatures are a far cry from today's High Arctic, where central Ellesmere Island experiences a mean annual temperature of −19 °C (−2 °F), a warm month mean temperature of about 6 °C (43 °F) and a cold month mean temperature of −38 °C (−36 °F) or colder.
      Despite the mild Eocene Arctic climate, the vertebrate fauna would have experienced months of total darkness and cooler temperatures during the winter. Recent isotopic work suggests that some mammals, including the hippo-like Coryphodon, were year-round residents in the High Arctic. Given that Gastornis was large (approaching
      2 metres (6.6 ft)) and flightless, it likely also was a year-round resident of the Arctic. In contrast, the volant Presbyornis might have been a seasonal migrant to the Arctic.


      Fossil content



      The formation has provided the following fossils:


      = Mammals

      =
      Primates
      Paromomyidae indet.
      Acreodi
      Pachyaena sp.
      Carnivora

      Dermoptera
      Plagiomene sp.
      Eutheria

      Ferae

      Glires

      Hyaenodonta
      Prolimnocyon sp.
      Leptictida
      Prodiacodon sp.
      Multituberculata

      Pantodonta
      Coryphodon pisuqti
      Perissodactyla


      = Birds

      =
      Anseriformes
      Presbyornis cf. pervetus
      Gastornithiformes
      Gastornis sp.


      = Reptiles

      =
      Turtles

      Crocodiles
      Allognathosuchus sp.
      Lizards

      Snakes
      Erycidae indet.


      = Amphibians

      =
      Caudata
      Piceoerpeton willwoodense


      = Fish

      =
      Teleostei
      Esox sp.
      Amiiformes

      Lepisosteiformes
      ?Lepisosteus sp.


      Correlations



      The formation has been correlated with the Allenby Formation and Kamloops Group of British Columbia, the Chickaloon Formation of Alaska, and with the Wishbone, Chuckanut and Iceberg Bay Formations. The upper Margaret Formation also has been correlated with the Buchanan Lake Formation of the Eastern Arctic Archipelago.


      See also




      References




      = Bibliography

      =
      Geology and climate
      West, Christopher K.; Greenwood, David R.; Reichgelt, Tammo; Lowe, Alexander J.; Vachon, Janelle M.; Basinger, James F. (2020), "Paleobotanical proxies for early Eocene climates and ecosystems in northern North America from middle to high latitudes", Climate of the Past, 16 (4): 1387–1410, Bibcode:2020CliPa..16.1387W, doi:10.5194/cp-16-1387-2020, retrieved 2020-09-05
      Sudermann, Markus; Galloway, Jennifer; Greenwood, David Robert; West, Christopher K.; Reinhardt, Lutz (2017), A palynological investigation of the Arctic late Paleocene–early Eocene Margaret Formation at Stenkul Fiord, Ellesmere Island, NU, Canada, Climate and Biotic Events of the Paleogene (CBEP 2017), p. 1, retrieved 2020-09-05
      Eberle, Jaelyn J.; Greenwood, David R. (2012), "Life at the top of the greenhouse Eocene world — A review of the Eocene flora and vertebrate fauna from Canada's High Arctic", GSA Bulletin, 124 (1–2): 3–23, Bibcode:2012GSAB..124....3E, doi:10.1130/B30571.1
      Schubert, Brian A.; Jahren, A. Hope; Eberle, Jaelyn J.; Sternberg, Leonel S.L.; Eberth, David A. (2012), "A summertime rainy season in the Arctic forests of the Eocene", Geology, 40 (6): 523–526, Bibcode:2012Geo....40..523S, doi:10.1130/G32856.1
      Reinhardt, Lutz; Andruleit, Harald; Estrada, Solveig; Henjes Kunst, Friedhelm; Piepjohn, Karsten (2010), Altered volcanic ashes in Paleocene/Eocene Eureka Sound Group sediments (Ellesmere Island, Arctic Canada) – new stratigraphic tie-points?, GeoCanada 2010 – Working with the Earth, pp. 1–2
      Paleontology
      Stidham, T. A.; Eberle, J. J. (2016), "The palaeobiology of high latitude birds from the early Eocene greenhouse of Ellesmere Island, Arctic Canada", Scientific Reports, 6: 1–8, Bibcode:2016NatSR...620912S, doi:10.1038/srep20912, PMC 4751439, PMID 26867798
      Beard, K. C.; Dawson, M. R. (2014), "Northernmost Global Record for Multituberculata from the Eocene of Ellesmere Island, Arctic Canada", Journal of Vertebrate Paleontology, 34 (6): 1476–1480, Bibcode:2014JVPal..34.1476B, doi:10.1080/02724634.2014.875032
      Dawson, M. R (2012), "Coryphodon, the northernmost Holarctic Paleogene pantodont (Mammalia), and its global wanderings", Swiss Journal of Palaeontology, 131 (1): 11–22, Bibcode:2012SwJP..131...11D, doi:10.1007/s13358-011-0028-1
      Gardner, J. D (2012), "Revision of Piceoerpeton MESZOELY (Caudata: Scapherpetontidae) and description of a new species from the late Maastrichtian and ? early Paleocene of western North America", Bulletin de la Société Géologique de France, 183 (6): 611–620, doi:10.2113/gssgfbull.183.6.611
      Alroy, J (2006), Synonymies and reidentifications of North American fossil vertebrates
      Eberle, J. J (2005), "A new 'tapir' from Ellesmere Island, Arctic Canada - Implications for northern high latitude palaeobiogeography and tapir palaeobiology", Palaeogeography, Palaeoclimatology, Palaeoecology, 227 (4): 311–322, Bibcode:2005PPP...227..311E, doi:10.1016/j.palaeo.2005.06.008
      Rose, K. D.; Eberle, J. J.; McKenna, M. C. (2004), "Arcticanodon dawsonae, a primitive new palaeanodont from the lower Eocene of Ellesmere Island, Canadian High Arctic", Canadian Journal of Earth Sciences, 41 (6): 757–763, Bibcode:2004CaJES..41..757R, doi:10.1139/e04-019
      Eberle, J. J.; McKenna, M. C. (2002), "Early Eocene Leptictida, Pantolesta, Creodonta, Carnivora, and Mesonychidae (Mammalia) from the Eureka Sound Group, Ellesmere Island, Nunavut", Canadian Journal of Earth Sciences, 39 (6): 899–910, Bibcode:2002CaJES..39..899E, doi:10.1139/e02-001
      Dawson, M. R (2001), "Early Eocene rodents (Mammalia) from the Eureka Sound Group of Ellesmere Island, Canada", Canadian Journal of Earth Sciences, 38 (7): 1107–1116, doi:10.1139/E01-010, S2CID 128408506
      Eberle, J. J (2001), "Early Eocene Leptictida, Pantolesta, Creodonta, Carnivora, and Cete from Ellesmere Island--Arctic Links to Europe and Asia", Journal of Vertebrate Paleontology, 21: 46A
      Dawson, M. R (1990), Canada's Missing Dimension
      Estes, R.; Hutchison, J. H. (1980), "Eocene lower vertebrates from Ellesmere Island, Canadian Arctic Archipelago", Palaeogeography, Palaeoclimatology, Palaeoecology, 30: 325–347, Bibcode:1980PPP....30..325E, doi:10.1016/0031-0182(80)90064-4
      McKenna, M. C (1980), "Eocene paleolatitude, climate, and mammals of Ellesmere Island", Palaeogeography, Palaeoclimatology, Palaeoecology, 30: 349–362, Bibcode:1980PPP....30..349M, doi:10.1016/0031-0182(80)90065-6
      West, R. M.; Dawson, M. R.; Hutchison, J. H. (1977), "Fossils from the Paleogene Eureka Sound Formation, N.W.T., Canada; occurrence, climatic and paleogeographic implications. In West, R. M. (ed), Paleontology and plate tectonics with special reference to the history of the Atlantic Ocean", Milwaukee Public Museum Contributions in Biology and Geology, 2: 77–93

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    Margaret Formation - Wikipedia

    The Margaret Formation is a geologic formation of the Eureka Sound Group in the Sverdrup Basin in Northwest Territories and Nunavut, Canada. The unit belonging to the Eureka Sound Group which crops out at Ellesmere Island preserves fossils dating back to the Early Eocene period, or Wasatchian in the NALMA classification. [1]

    Palynostratigraphy of the lower Paleogene Margaret Formation at …

    03 Jul 2021 · The upper Paleocene to lower Eocene Margaret Formation exposed at Stenkul Fiord on southern Ellesmere Island, Nunavut, Canada, represents a nearly continuous terrestrial succession of microfossil-rich clastic sediments and coal.

    Margaret Formation - Wikiwand

    The Margaret Formation is a geologic formation of the Eureka Sound Group in the Sverdrup Basin in Northwest Territories and Nunavut, Canada. The unit belonging ...

    Palynostratigraphy of the Lower Paleogene Margaret Formation …

    06 Agu 2021 · The upper Paleocene to lower Eocene Margaret Formation exposed at Stenkul Fiord on southern Ellesmere Island, Nunavut, Canada, represents a nearly continuous terrestrial succession of microfossilrich clastic sediments and coal.

    Paleogene sedimentation and Eurekan deformation in the …

    14 Jun 2019 · Field studies and interpretative mapping of the area southeast of Stenkul Fiord (Ellesmere Island) revealed that the Margaret Formation clastic deposits consist of at least four sedimentary units (Units 1–4) separated by unconformities.

    The palaeobiology of high latitude birds from the early Eocene ...

    12 Feb 2016 · The Margaret Formation comprises coarsening-upward sequences of interbedded cross-bedded sandstone, siltstone, mudstone and coal that are interpreted as proximal delta-front to delta-plain ...

    (PDF) Palynostratigraphy of the lower Paleogene Margaret Formation …

    27 Jan 2021 · The upper Paleocene to lower Eocene Margaret Formation exposed at Stenkul Fiord on southern Ellesmere Island, Nunavut, Canada, represents a nearly continuous terrestrial succession of...

    Arctic Canadian Paleocene and Eocene fossil floras and paleoclimates ...

    Examples of the fossil leaf morphotypes from the Margaret Formation, Stenkul Fiord, Ellesmere island. From West et al. (2015). The project was initially based on the extensive collections of Jim Basinger and his associates (including Greenwood) housed at the University of Saskatchewan.

    Palynostratigraphy of the lower Paleogene Margaret Formation at Stenkul ...

    The upper Paleocene to lower Eocene Margaret Formation exposed at Stenkul Fiord on southern Ellesmere Island, Nunavut, Canada, represents a nearly continuous terrestrial succession of microfossil-rich clastic sediments and coal.

    Additions to the Eocene Perissodactyla of the Margaret Formation ...

    09 Jan 2015 · We describe early Eocene (Wasatchian) occurrences of the isectolophid Homogalax, tapiroids Heptodon posticus, Heptodon cf. H. posticus, and Heptodon sp., as well as early middle Eocene (Bridgerian) fossils of the brontothere Palaeosyops from localities in the Margaret Formation of the Eureka Sound Group on Ellesmere Island, Nunavut, Arctic Canada.