- Source: Clavatulidae
Clavatulidae is a taxonomic family of sea snails, marine gastropod mollusks in the superfamily Conoidea. The family is not well differentiated morphologically.
Clavatulidae was raised, based on cladistic analysis, from subfamily to the family level by Rosenberg in 1998. It is no longer regarded as a subfamily of Turridae by several malacologists (Kantor, Sysoev). This family has no subfamilies.
General characteristics
This family consists of species with a medium-sized to rather large, fusiform shell. The oblong, pointed spire is rather high. The aperture is oval and mostly white. The operculum has a medio-lateral nucleus. The siphonal canal varies between rather short (e.g. Pusionella compacta) to moderately long and slightly incurved (e.g. Fusiturris undatiruga). The anal sinus varies from very shallow to rather deep. The outer lip can be slightly incurved and serrated on its side. The subsutural ramp is usually well developed. The sculpture of the shell in this family shows various forms, going from a rather smooth surface (e.g. Gemmuloborsonia colorata) to being finely ribbed longitudinally and striated transversally. The stenoglossan radula has the formula 1-(1-R-1)-1
Genera
Genera in the family Clavatulidae include:
Benthoclionella Kilburn, 1974
Caliendrula Kilburn, 1985
Clavatula Lamarck, 1801
Clionella Gray, 1847
† Granulatocincta Harzhauser, Landau & R. Janssen, 2022
† Hemisurcula Casey, 1904
Makiyamaia Kuroda, 1961
† Megaclavatula Harzhauser, Landau & R. Janssen, 2022
† Neoperrona Harzhauser, Landau & R. Janssen, 2022
† Orthosurcula T. L. Casey, 1904
Pagodaturris Kantor, Fedosov & Puillandre, 2018
Paraclavatula Kantor, Horro, Rolán & Puillandre, 2018
Perrona Schumacher, 1817
Pusionella Gray, 1847
Scaevatula Gofas, 1990
† Striopusionella Harzhauser, Landau & R. Janssen, 2022
Tomellana Wenz, 1943
Toxiclionella Powell, 1966
† Trachelochetus Cossmann, 1889
Trachydrillia Nolf & Swinnen, 2010
Turricula Schumacher, 1817
Genera moved to other families
Danilacarina Bozzetti, 1997, moved to Cochlespiridae
Iwaoa Kuroda, 1953, moved to Horaiclavidae
Genera brought into synonymy
Melatoma Swainson, 1840: synonym of Clionella Gray, 1847
Netrum Philippi, 1850: synonym of Pusionella Gray, 1847
Surcula H. Adams & A. Adams, 1853: synonym of Turricula Schumacher, 1817
Tomella Swainson, 1840: synonym of Tomellana Wenz, 1943
†Trachelochetus Cossmann, 1889: synonym of †Clavatula (Trachelochetus) Cossmann, 1889 represented as Clavatula Lamarck, 1801
Tyrrhenoturris Coen, 1929: synonym of Fusiturris Thiele, 1929
Ecology
Species from family Clavatulidae are omnivores, predators and scavengers.
References
External links
Worldwide Mollusc Species Data Base: family Clavatulidae