- Source: Notocupes
Notocupes is an extinct genus of medium-sized archostematan beetles from the Mesozoic Era of Eurasia, including over 50 described species. Historically, the genus was classified as a member of the family Ommatidae, but the presence of characters such as the horizontal mandibular cutting edge, separated procoxae and overlapping abdominal sternites indicate that the genus may have a closer affinity with the family Cupedidae. Notocupes is considered to be a junior synonym of Zygadenia by Kirejtshuk (2020), but other researchers suggest to reserve the genus Zygadenia as a form taxon for isolated elytra that probably belong to the genus Notocupes, while retaining Notocupes as a valid genus for complete body fossils. Most species of Notocupes were described from compression fossils. An additional three species were described from Cenomanian-aged Burmese amber, which were treated as a separate genus, Echinocups, by Kirejtshuk (2020), but Li et al. (2023) consider Echinocups to be a junior synonym of Notocupes. Notocupes has a flattened body, which may suggest that it occupied narrow habitats, such as living under bark. Some species had serrated/spined margins of the carapace, which may have served as a defense against predators, or served as camouflage to resemble bark.
Strelnikova & Yan (2023) advise splitting up the genus Notocupes into four genera: Notocupes, Rhabdocupes, Conexicoxa and the new genus Brachilatus.
Triassic species
= Ladinian (242 Ma to ~237 Ma)
=Tongchuan Formation, China
N. shiluoensis (Hong, 1984) = Chengdecupes shiluoensis
= Carnian (237 Ma to 227 Ma)
=Koldzat Formation, Kazakhstan
N. oxypygus Ponomarenko, 1969
Madygen Formation, Kyrgyzstan
N. laticella Ponomarenko, 1969
N. rostratus Ponomarenko, 1969
N. tenuis Ponomarenko, 1969
Jurassic species
= Hettangian (201.3 Ma to 199.3 Ma)
=Dzhil Formation, Kyrgyzstan
N. cellulosus Ponomarenko, 1969
N. issykkulensis Ponomarenko, 1969
N. kirghizicus Ponomarenko, 1969
N. latus Ponomarenko, 1969
N. sogutensis Ponomarenko, 1969
Guanyintan Formation, China
N. homorus (Lin, 1986) = Conexicoxa homora
= Toarcian (182.7 Ma to 174.1 Ma)
=Guanyintan Formation, China
N. lini Ponomarenko et al., 2012
Sulyukta Formation, Kyrgyzstan
N. crassus Ponomarenko, 1969
= Bajocian (170.3 Ma to 168.3 Ma)
=Bakhar Formation, Mongolia
N. brachycephalus Ponomarenko, 1994
N. exiguus Ponomarenko, 1994
N. longicollis Ponomarenko, 1994
= Callovian (166.1 Ma to 163.5 Ma)
=Haifanggou Formation, China
N. daohugouensis Li and Cai in Lin et al., 2023
N. dischides Zhang, 1986
N. jurassicus (Hong, 1983) = Chengdecupes jurassicus
N. pingi Ponomarenko and Ren, 2010
N. robustus Li and Cai in Lin et al., 2023
N. spinosus Li and Cai in Lin et al., 2023
Tyumen Formation, Russia
N. patulus Ponomarenko, 1985
= Oxfordian (163.5 Ma to 157.3 Ma)
=Karabastau Formation, Kazakhstan
N. foersteri Ponomarenko, 1971
N. lapidarius Ponomarenko, 1968
N. nigromonticola Ponomarenko, 1968
N. picturatus Ponomarenko, 1964
N. pulcher Ponomarenko, 1968
= Tithonian (152.1 Ma To 145.0 Ma)
=Shar Teeg, Mongolia
N. brachycephalus Ponomarenko, 1994
N. exiguus Ponomarenko, 1994
Solnhofen, Germany
N. reticulatus (Oppenheim, 1888) = Procarabus reticulatus
N. tripartitus (Oppenheim, 1888) = Procarabus tripartitus
Cretaceous species
= Hauterivian (132.9 Ma to 129.4 Ma)
=Dabeigou Formation, China
N. lentus (Ren, 1995) = Tetraphalerus lentus
= Barremian (129.4 Ma to 125.0 Ma)
=Jianshangou Formation, China
N. laetus (Lin, 1976)
La Huérguina Formation, Las Hoyas, Spain
N. diazromerali (Soriano & Delclòs, 2006) = Zygadenia diazromerali
N. longicoxa (Soriano & Delclòs, 2006) = Zygadenia longicoxa
N. siniestri (Soriano & Delclòs, 2006) = Zygadenia siniestri
N. viridis (Soriano & Delclòs, 2006) = Zygadenia viridis
La Pedrera de Rúbies Formation, Las Hoyas, Spain
N. martinclosas (Soriano & Delclòs, 2006) = Zygadenia martinclosas
N. oculatus (Soriano & Delclòs, 2006) = Zygadenia oculata
Ulan-Argalant Formation, Mongolia
N. mongolicus Ponomarenko, 1994
= Aptian (125.0 to ~113.0 Ma)
=Argun Formation, Russia
N. semen (Ponomarenko, 2000) = Zygadenia semen
N. sibiricus (Ponomarenko, 2000) = Zygadenia sibirica
Baojiatun Formation, China
N. baojiatunensis (Hong, 1992) = Chengdecupes baojiatunensis
Dzun-Bain Formation, Mongolia
N. dundulaensis Ponomarenko, 1994
N. elegans Ponomarenko, 1994
Jehol Group, China
N. validus (Lin, 1976) = Sinocupes validus
Khasurty Formation, Russia
N. khasurtyiensis Strelnikova, 2019
Laiyang Formation, China
N. laiyangensis (Hong, 1990) = Forticupes laiyangensis
N. ludongensis Wang and Liu, 1996
N. tuanwangensis (Hong, 1990) = Picticupes tuanwangensis
Shahai Formation, China
N. kezuoensis (Hong, 1987) = Chengdecupes kezuoensis
Yixian Formation, China
N. alienus (Tan and Ren, 2006) = Ovatocupes alienus
N. cyclodontus (Tan et al., 2006) = Amblomma cyclodonta
N. epicharis (Tan et al., 2005) = Amblomma epicharis
N. eumeurus (Tan et al., 2006) = Amblomma eumeura
N. minisculus (Tan et al., 2006) = Amblomma miniscula
N. porrectus (Tan et al., 2006) = Amblomma porrecta
N. protensus (Tan et al., 2006) = Amblomma protensa
N. psilatus (Tan et al., 2005) = Amblomma psilata
N. rudis (Tan et al., 2006) = Amblomma rudis
N. stabilis (Tan et al., 2006) = Amblomma stabilis
Zaza Formation, Russia
N. caudatus Ponomarenko, 1966
N. excellens Ponomarenko, 1966
N. vitimensis Ponomarenko, 1966
= Albian (~113.0 to 100.5 Ma)
=Jinju Formation, South Korea
N. premeris Lee et al., 2022
= Cenomanian (100.5 to 93.9 Ma)
=Burmese amber, Myanmar
N. denticollis Jiang et al., 2020 (sometimes placed in separate genus Echinocups)
N. neli Tihelka et al., 2019 (sometimes placed in separate genus Echinocups)
N. ohmkuhnlei Jarzembowski et al., 2019 (sometimes placed in separate genus Echinocups)
= Turonian (93.9 to 89.8 Ma)
=Emanra Formation, Russia
N. khetanensis Ponomarenko in Cromov et al., 1993
Kzyl-Zhar Locality, Kazakhstan
N. caducus Ponomarenko, 1969
References
Kata Kunci Pencarian:
- Notocupes
- Cupedidae
- Ommatidae
- Echinocups
- Rhabdocupes
- Notocupoides
- Archostemata
- Eurydictyon
- Zygadenia
- Telephone-pole beetle
