- Source: Portunoidea
Portunoidea is a superfamily of crabs that includes the family Portunidae, the swimming crabs. Which other crab families are also placed here is a matter of some contention, and may be revised following molecular phylogenetic analyses.
Description
Their rather flat and smooth carapace is usually wider than long and of hexagonal, subhexagonal, rectangular, or transversely ovate shape. It is usually widest between the hindmost spines of the forward rim; there may be up to 9 pairs of these spines, with a few smaller ones right above the head, but they are missing altogether in some species.
In some, the first maxilliped's endopod is lobed, forming the characteristic portunid lobe. The chelipeds are usually robust, and in some the last pereiopod pair has ovate dactyls. The sutures of the sternum between segments 4 to 8 are usually incomplete, and in the Portunidae, the eighth sternite is usually visible if seen from below and has a penial groove.
In males, the abdominal somites are either all free or the third to fifth are fused, often retaining the sutures though. The first gonopod is strongly curved, with a swollen and strongly hooked base.
Classification
Portunoidea are close relatives of the Xanthoidea, and the families Hexapodidae and Mathildellidae, usually included there, are sometimes placed in the Portunoidea, while the deep-sea crabs (Geryonidae) are usually placed in the Portunoidea but sometimes in the Xanthoidea. All Portunoidea live in the ocean, although the family Trichodactylidae, sometimes included here, live in fresh water.
According to the latest synopsis, there are fourteen families in the superfamily Portunoidea, five of which are extinct:
Brusiniidae Števčić, 1991
†Carcineretidae Beurlen, 1930
Carcinidae MacLeay, 1838
†Eogeryonidae Ossó, 2021
Geryonidae Colosi, 1924
†Lithophylacidae Van Straelen, 1936
†Longusorbiidae Karasawa, Schweitzer & Feldmann, 2008
Nautilocorystidae Ortmann, 1893
Ovalipidae Spiridonov, Neretina & Schepetov, 2014
Pirimelidae Alcock, 1899
Polybiidae Ortmann, 1893
Portunidae Rafinesque, 1815
†Psammocarcinidae Beurlen, 1930
Thiidae Dana, 1852
It also contains the extinct genus Dinocarcinus.
References
Media related to Portunoidea at Wikimedia Commons