• Source: Timeline of dromaeosaurid research

This timeline of dromaeosaurid research is a chronological listing of events in the history of paleontology focused on the dromaeosaurids, a group of sickle-clawed, bird-like theropod dinosaurs including animals like Velociraptor. Since the Native Americans of Montana used the sediments of the Cloverly Formation to produce pigments, they may have encountered remains of the dromaeosaurid Deinonychus hundreds of years before these fossils came to the attention of formally trained scientists.
In 1922 Matthew and Brown named the new genus and species Dromaeosaurus albertensis, considering it a new type within the family Deinodontidae, a now-defunct family name that once applied to the tyrannosaurs. Not long after, Velociraptor was discovered in Mongolia by the Central Asiatic Expedition. Dromaeosaur research was fairly quiet until the 1960s, when John Ostrom described the new genus and species Deinonychus antirrhopus. This discovery played a major role in setting off the Dinosaur Renaissance because Deinonychus was obviously a vigorous, active animal, and exhibited characteristics linking it to the origin of birds. As such it brought support for controversial reinterpretations of dinosaurs as warm-blooded and ancestral to birds. Its distinct nature and similarity to Dromaeosaurus led Ostrom to follow Edwin Colbert and Dale Russel's suggestion that the Dromaeosaurinae be regarded as its own family separate from the Deinodontidae.
After Ostrom's initial research on Deinonychus, evidence continued to mount for a close evolutionary relationship between dromaeosaurids and birds. The dromaeosaurid Sinornithosaurus milennii, described in 1999 by Xu, Wang, and Wu, is a notable example as the fine-grained Chinese limestone from which it was collected preserved its life covering of feathers. Discoveries of feathered dromaeosaurids continued into the 2000s. Xu, Zhou, and Wang named the new genus Microraptor in 2000. Three years later, Xu and others would report a new species in this genus that exhibited a bizarre "four winged" body plan with long pennaceous flight feathers on both its front and hind limbs.




Prescientific



The Crow people and other Native American groups inhabiting Montana used to use rocks from the Cloverly Formation to make red pigments. Since the red pigments are richest in the same layers of the formation that preserve dinosaur fossils, it is likely that Native Americans encountered Deinonychus fossils long before scientifically trained paleontologists.




19th century






= 1880s

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1887

Harry Govier Seeley described the new species Ornithodesmus cluniculus.




20th century






= 1910s

=
1914

Barnum Brown collected a partial skull and foot in Late Cretaceous rocks that would later serve as the type specimen of Dromaeosaurus albertensis.




= 1920s

=

1922

Matthew and Brown described the new genus and species Dromaeosaurus albertensis and named the Dromaeosauridae.
1924

Henry Fairfield Osborn described the new genus and species Velociraptor mongoliensis.
1926

Matthew and Brown first observed that the braincase of dromaeosaurids were large for reptiles of their body size.




= 1960s

=

1969

John Ostrom described the new genus and species Deinonychus antirrhopus. This description provided the scientific literature with its first detailed information about the anatomy of the dromaeosaurid body, as previous work focused on the skull. He proposed that Deinonychus killed its prey with the enlarged, sharply curved claws on the second toe of its feet.
Ostrom further described the anatomy of Deinonychus antirrhopus, providing further details about the anatomy of its body. Ostrom proposed that Deinonychus only walked on two of its toes.




= 1970s

=

1972

Zofia Kielan-Jaworowska and Rinchen Barsbold reported the associated remains of a Velociraptor and Protoceratops apparently killed and preserved while fighting.
1973

Ostrom first discussed the apparent close evolutionary relationship between Deinonychus antirrhopus and birds.
1975

Ostrom published further arguments building a case for a close evolutionary relationship between dromaeosaurids and birds.

1976

Ostrom published further arguments building a case for a close evolutionary relationship between dromaeosaurids and birds.
1978

Sues described the new genus and species Saurornitholestes langstoni.
1979

Powell described the new genus and species Unquillosaurus ceibalii.




= 1980s

=

1981

Jensen described the new genus and species Palaeopteryx thomsoni.
1982

Osmolska described the new genus and species Hulsanpes perlei.
1983

Rinchen Barsbold divided the Dromaeosauridae into two subfamilies; the Dromaeosaurinae and Velociraptorinae.
Barsbold described the new genus and species Adasaurus mongoliensis. Unlike most dromaeosaurs, famous for the "killing claws" on their second toe, the second toe claw of A. mongoliensis was relatively small and not sharply curved. It is similar to those of troodontids.
1985

Ostrom published further arguments building a case for a close evolutionary relationship between dromaeosaurids and birds.
1986

Gauthier found strong support for Ostrom's hypothesis that dromaeosaurids were closely related to birds in "a pioneering and thorough analysis".




= 1990s

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1990

Ostrom published further arguments building a case for a close evolutionary relationship between dromaeosaurids and birds.
1993

Kirkland, Burge, and Gaston described the new genus and species Utahraptor ostrommaysi.
Osmolska challenged Kielan-Jaworowska and Barsbold's 1972 interpretation of an associated skeleton of Velociraptor and Protoceratops as being of two animals killed and preserved while locked in mortal combat. She reinterpreted the association as a Velociraptor that had been killed while scavenging an already dead Protoceratops.

1994

Ostrom speculated that Deinonychus hunted in packs to subdue prey larger than itself based on an association between several Deinonychus skeletons and a specimen of the ornithopod Tenontosaurus.
1995

Currie followed Barsbold's division of Dromaeosauridae into Dromaeosaurinae and Velociraptorinae. However, he proposed differing definitions for these subfamilies and argued that they had different members than were included under Barsbold's classification scheme.
Maxwell and Ostrom reported on the association of Deinonychus teeth and Tenontosaurus remains and argued that these provided further support for the hypothesis that Deinonychus was a pack hunter.
Unwin and others followed the fighting-to-the-death interpretation of the fighting dinosaurs specimen.

1997

Novas and Puerta described the new genus and species Unenlagia comahuensis.
Chatterjee argued that dromaeosaurids lived in trees based on the anatomy of their hands and the backward orientation of the pubic bones in their pelvis.
Reid examined histological sections of the bones of Saurornitholestes to see whether or not they were consistent with the idea that dromaeosaurids were warm blooded. However, the results of his study were inconclusive.
Ruben and others attempted to reconstruct the way air would flow through the snouts of dromaeosaurids during respiration to see whether or not it was consistent with the airflow patterns of modern cold or warm-blooded animals. They found that the probable course of airflow through the snout of a living, breathing dromaeosaurid would have been more consistent with that of a cold blooded animal. However, the anatomical reconstructions the researchers used to draw these conclusions have been criticized for positioning the choana too far forward and for excluding the animals' secondary palate.
1998

Le Leouff and Buffetaut described the new genus and species Variraptor mechinorum.

D. L. Brinkman and others reinterpreted the association between several Deinonychus specimens and a Tenontosaurus as preserving the aftermath of an ancient feeding frenzy rather than as evidence of pack hunting behavior in Deinonychus as argued by Ostrom.
1999

Clark and others argued that it was impossible to know whether or not Velociraptor was a predator of Protoceratops based on the famous "fighting dinosaurs" specimen, since this was only one specimen.
Altangerel Perle, Norell and Clark described the new genus and species Achillobator giganticus.
Xu, Wang, and Wu described the new genus and species Sinornithosaurus millenii. The specimen actually preserved the remains of the feathers that covered the animal in life. The presence of a feathery covering is consistent with the idea that dromaeosaurids were warm-blooded. They also performed a cladistic analysis of the Dromaeosauridae, finding more support for dividing the family into Dromaeosaurinae and Velociraptorinae. The researchers found both groups to be monophyletic clades, with Sinornithosaurus milenii the sister group to both, and therefore the most primitive known dromaeosaurid.




21st century






= 2000s

=
2000

Burnham and others described the new genus and species Bambiraptor feinbergorum.
Xu, Zhou and Wang described the new genus and species Microraptor zhaoianus.
Allain and Taquet described the new genus and species Pyroraptor olympius.

2001

A. R. Jacobsen published a description of a dentary referred to Saurornitholestes with tooth marks. The specimen was preserved in the Dinosaur Park Formation. Although a specific identification cannot be made, the shape of the preserved serration marks implicate a juvenile individual of one of the formation's tyrannosaurids, like Gorgosaurus, Daspletosaurus, or Aublysodon. All of the marks on the jawbone seem to have been left by the same animal because the serration marks all share the same morphology.
Kevin Padian, Ji Qiang and Ji Shu-an published a review of known feathered dinosaurs and their implications for the origin of flight. The authors observed that many aspects of the distribution of feather homologues in the dinosaur family tree met the expectations of earlier phylogenetic hypotheses, including a gradual transition from primitive filaments in Sinosauropteryx to the shared filaments and "rudimentary" true feathers in Caudipteryx and Protarchaeopteryx, to flight feathers in Archaeopteryx. However, they also noted, only filamentous protofeathers had been observed in dromaeosaurs like Sinornithosaurus, when Caudipteryx-style true feathers would be expected due to the family's phylogenetic position. Also noted were the lack of known integumentary structures among troodontids. The authors proposed that either science's understanding of coelurosaur evolution is faulty or feathers evolved multiple times.

2002

Hwang and others performed a cladistic analysis of the Dromaeosauridae.
Xu and others performed a cladistic analysis of the Dromaeosauridae.

Czerkas and others described the new genus and species Cryptovolans pauli.
2003

Christiansen and Bonde described the new genus and species Dromaeosauroides bornholmensis.
2004

Currie and Varricchio described the new genus and species Atrociraptor marshalli.

Xu and Wang described the new genus and species Graciliraptor lujiatunensis.
2005

Makovicky, Apesteguía and Agnolin described the new genus and species Buitreraptor gonzalezorum.
Novas and Pol described the new genus and species Neuquenraptor argentinus.
2006

Norell and others described the new genus and species Tsaagan mangas.
2007

Lü and others described the new genus and species Luanchuanraptor henanensis.

Turner and others described the new genus and species Mahakala omnogovae.
A. S. Turner, S. H. Hwang, and M. A. Norell described the new genus and species Shanag ashile.
2008

Novas et al. described the new genus and species Austroraptor cabazai.
2009

Longrich and Currie described the new genus and species Hesperonychus elizabethae.




= 2010s

=
2010

Xu and others described the new genus and species Linheraptor exquisitus.
Zheng and others described the new genus and species Tianyuraptor ostromi.
2011

Porfiri, Calvo and Santos described the new genus and species Pamparaptor micros.
2012

Gong and others described the new species Microraptor hanqingi.

Senter and others described the new genus and species Yurgovuchia doellingi.
2013

D. C. Evans, D. W. Larson, and P. J. Currie described the new genus and species Acheroraptor temertyorum.
2014

Han and others described the new genus and species Changyuraptor yangi.
2015

Jasinski described the new species Saurornitholestes sullivani.
Lü and Brusatte described the new genus and species Zhenyuanlong suni.
DePalma et al. described the new genus and species Dakotaraptor steini.

Bell and Currie described the new genus and species Boreonykus certekorum.
2017

Xu and others described the new genus and species Zhongjianosaurus yangi.
Cau and others described the new genus and species Halszkaraptor escuilliei.
2019

An ungual phalanx of a dromaeosaurid theropod is described from the Blagoveshchensk area (Russia) by Bolotskii, Bolotskii & Sorokin (2019).




See also


History of paleontology
Timeline of paleontology




Footnotes






References






External links


Media related to Dromaeosauridae at Wikimedia Commons

Kata Kunci Pencarian:

• Timeline of dromaeosaurid research
• Dromaeosauridae
• Timeline of troodontid research
• Utahraptor
• Unenlagiinae
• Velociraptor
• Fighting Dinosaurs
• Neuquenraptor
• Austroraptor
• Itemirus